Fig. 1 Schematic view of calculation region

Fig. 2 Concept of ecosystem model

The material cycle process in each ecosystem model consists of physical process, such as transfer and diffusion, and biological processll) as follows;

・Phytoplankton (P) [mgC/m³]

dP/dt = (Transfer & Diffusion) + (Nutrient consumption) - (Secretion) - (Natural death) - (Predation by zooplanlcton) - (Filtering by benthos) - (Sticking to seaweed) - (Sticking to breakwater) = (Transfer & Diffusion) + V1(tc,N)- V2(I(t))・ P - β (P)・V1(tc,N)・V2(I(t))・P - α ・P - P/(P+D)・g(P,D)・Z - P/(P+Z+D)・f(O,tc)・B - P/(P+Z+D)・ τ・S- rs・(1-k)・P (1)

・Zooplanlcton (Z) [mgC/m³]

dZ/dt = (Transfer & Diffusion) + (predation) - (Evacuation) - (Excretion) - (Natural death) - (Filtering by benthos) - (Sticking to seaweed) - (Sticking to breakwater) = (Transfer & Diffusion) + g(P,D)・Z - (1-μ )・g(P,D)・Z - (μ - ν )・g(P, D)・Z - δ・Z - Z/(P+Z+D).f(O,tc)・B - Z/(P+Z+D)・τ・S- rs・(1-k)・Z (2)

・Detritus (D) [mgC/m³]

dD/dt = (Transfer & Diffusion) + (Secretion of phyto- plankton)+ (Secretion of Seaweed) + (Natural death of phyto- plankton) + (Natural death of zooplankton) + (Natural death of seaweed) + (Evacuation of zooplankton) - (Predation by zooplanicton) - (Decomposition) - (Filtering by benthos)- (Sticking to seaweed) - (Sticking to brealcwater) = (Transfer & Diffusion) + β (P)・V1(tc,N)・ V2(1(t))・P + β(S)・ V1'(tc, N)・V2'（I(t))・S+α・P+δ・Z+α S・S +(1-μ)・g(P,D)・Z- D/(P+D)・g(P,D)・Z - φ ・D - D/(P+Z+D)・f(O,tc)・B - D/(P+ Z+D)・ τ ・S-rs,(1-k)・D (3)

・Nutrients (N) [mgN/m³]

dN/dt = (Transfer & Diffusion) + (Excretion of zooplanicton) + (Excretion of benthos) + (Decomposition of detritus) - (Consumnption by phytoplankton) - (Consumption by seaweed) - (Sticking to seaweed) - (Sticking to brealcwater) = (Transfer & Diffusion) + (μ - ν )・g(P,D)・Z+ (μｂ-νｂ)・ f(O,tc) B+ φ・D - V1(tc,N)・ V2(I(t)）・ P - V1'(tc,N). V2'(I(t)). S - N/(P+ Z+D)・τ・S-rs・(l-k)-N (4)

・Benthos (B) [mgC/m³]

dB/dt = (Phytoplankton filtering) + (Zooplanicton filtering) + (Detritus filtering) - (Evacuation) - (Excretion) - (Natural death) = f(O,tc)・L（B)・B - (i-μ b)・f(O,tc)・L(B).B (μb - νb)・f(O,tc)・L(B）B- δb・B (5)

・ Seaweeds (S) [mgC/m³]

dS/dt = (Nutrient consumption) - (Secretion) - (Natural death) ＝ V1'(tc,N)・ V2'(I(t))・ '(S)・S - (β (S)・ V1'(tc,N)・ V2'(I(t))・ L' (S)・S-αｓa・S (6)

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